Type Specimens of Dung Flies (Diptera, Scathophagidae) in the Collection of the Zoological Institute of the Russian Academy of Sciences in St. Petersburg

Abstract

A catalogue of type specimens (holotypes, paratypes, lectotypes, and paralectotypes) of species of the family Scathophagidae (Diptera) described by Th. Becker, I.A. Portschinsky, A.A. Stackelberg, K.B. Gorodkov, A.L. Ozerov, and M.G. Krivosheina, which are deposited in the collection of the Zoological Institute of the Russian Academy of Sciences in St. Petersburg, Russia, is presented. Photographs of the specimens and labels of the holotypes and lectotypes are given.

All the photographs were taken by N.M. Paramonov using a Canon EOS 800D digital camera with a 65 mm MP-E lens. The images were processed using Helicon Focus 6 software.

The names of the species are listed in alphabetic order and followed by the quoted labels of the type specimens; the current name of the species is given when it differs from the original combination. The labels of the primary types are quoted verbatim, and those of the remaining specimens are given in their complete current or original form in order to make their contents as understandable as possible. Circles of golden paper are pinned under some specimens to designate their belonging to the type series.

The following abbreviations of the names of the institutes and museums are used below:

MNB, Museum für Naturkunde Berlin / Natural History Museum, Berlin, Germany;

ZIN, Zoological Institute of the Russian Academy of Sciences, St. Petersburg, Russia;

ZMMSU, Zoological Museum of M.V. Lomonosov Moscow State University, Moscow, Russia.

Species Described by Th. Becker

albidohirta Becker, 1907 (Becker, 1907 : 254 (Scatophaga)) (Fig. 1). The species was described based on 8 specimens. Lectotype, male: “[Kitai] Kurlyk, Baingol, vost[ochnyi] Tsaidam 28.V.[18]95 (Rob[orovskii] Kozlov)” [in Cyrillic, = China, Kurlyk, Baingol, East Qaidam, 28.V.[18]95 (Roborovsky, Kozlov)] [Ozerov and Krivosheina (2011) erroneously cited the date as 21.V], “Syntypus Scatophaga albidohirta Becker, 1907,” handwritten label “A. Ozerov. 20.11.2010” [in Cyrillic], “Lectotypus, ♂ Scatophaga albidohirta Becker, 1907, designation by A.L. Ozerov, 2010,” designated by A.L. Ozerov (Ozerov and Krivosheina, 2011). The inventory number is INS_DIP_0001151. The abdomen of the lectotype is cut off and placed together with the genitalia in a vial pinned under the specimen. Both middle legs are missing; the remains of one middle leg are glued onto the locality label (a fragment of the tibia is visible in the photo); the left hind tarsus and tibia are missing; only the 1st tarsomere is preserved in the right fore tarsus; both wings are partly damaged.

Fig. 1.

Scathophaga albidohirta Becker, 1907, lectotype. Scale bar 1 mm.

Paralectotypes, 2 females: “[Kitai] r[eka] Orogyn Syrtyn yu[zhnee] Nan’shanya, Gobi 3–20.VII.[18]95 (Rob[orovskii], Kozlov)” [in Cyrillic, = China, Orogyn River, Syrtyn S of Nanshan, Gobi, 3–20.VII.[18]95, Roborovsky, Kozlov]; 1 male: “[Kitai] oz[ero] Iche – r[eka] Orogyn, s[evernyi] Tsaidam”, ts[entral’naya] Gobi 1–3.VII.[18]95 (Rob[orovskii], Kozlov)” [in Cyrillic, = China, Lake Iche, Orogyn River, North Qaidam, Central Gobi, 1–3.VII.[18]95, Roborovsky, Kozlov]. The latter specimen was not examined by A.L. Ozerov and was not mentioned in his paper written together with M.G. Krivosheina (Ozerov and Krivosheina, 2011).

The male labeled “Kurlyk, Baingol, East Qaidam, 16–24.V.[18]95 (Roborovsky, Kozlov)” [in Cyrillic] was identified by A.L. Ozerov (Ozerov and Krivosheina, 2011) as Scathophaga mihalyii (Šifner, 1975). Two females with labels “Kara Tube, W of Khami, Gobi 15.IX.[18]95 (Roborovsky, Kozlov)” [in Cyrillic; Th. Becker commented on this label as “Gobi im Chines. Turkestan”] and “Bomyn (Ichegyn) River, Northeast Qaidam, Gobi, end of VI.[18]95 (Roborovsky, Kozlov)” [in Cyrillic] have uniformly yellow tarsi without yellow rings on the tarsomeres and definitely do not belong to Scathophaga albidohirta. These two specimens cannot be identified to species, and they were therefore labeled by A.L. Ozerov as Scathophaga sp. (Ozerov and Krivosheina, 2011).

The eighth syntype, male: “oz[ero] Iche – r[eka] Orogyn, s[evernyi] Tsaidam ts[entral’naya] Gobi (Rob[orovskii], Kozlov) 3.VII.[18]95” [in Cyrillic, = Lake Iche, Orogyn River, North Qaidam, Central Gobi (Roborovsky, Kozlov), 3.VII.[18]95], “Gobi, Asien VII. 51896 [?1895],” “albidihirta Bec[ker].” The latter two labels, one of them with the specific epithet misspelled, were written by Th. Becker (according to Dr. J. Ziegler, pers. comm. to A.L. Ozerov; see Ozerov and Krivosheina, 2011). This specimen, deposited at MNB, was examined by A.L. Ozerov in 2012 and also designated as a paralectotype (Ozerov and Krivosheina, 2011).

arctica Becker, 1897 (Becker, 1897 : 398 (Scatophaga)) (Fig. 2). The species was described based on 4 specimens. Lectotype, male, designated by A.L. Ozerov (Ozerov and Krivosheina, 2013): “Mal[ye] Karmakuly, Novaya Zemlya. 31.VII.[18]96 (Yakobson)” [in Cyrillic, = Malye Karmakuly, Novaya Zemlya, 31.VII.[18]96 (Jacobson)], “Scatophaga arctica Beck.,” “Scatophaga apicalis Curtis [in Ross, 1835] ♂ Gorodkov det. [19]73” (Gorodkov, 1986), “Lectotypus, ♂ Scatophaga arctica Becker, 1897: 398 designation by A.L. Ozerov, 2011,” “Scathophaga multisetosa (Holmgren, [1883]) det. A.L. Ozerov, 2013.” The inventory number is INS_DIP_0001152. The abdomen is cut off and placed together with the genitalia in a vial pinned under the specimen; otherwise the lectotype is well preserved.

Fig. 2.

Scathophaga arctica Becker, 1897, lectotype. Scale bar 1 mm.

Paralectotype, female: “stoyanka u g[ryady] Chernysheva, Nov[aya] Zem[lya] 4.VIII.[18]96 (Yakobson)” [in Cyrillic, = camp near Chernysheva Ridge, Novaya Zemlya, 4.VIII.[18]96 (Jacobson)]. One more paralectotype could not be located. The current name is Scathophaga apicalis (Curtis in Ross, 1835) (Ozerov and Krivosheina, 2023).

infumatum Becker, 1907 (Becker, 1907 : 256 (Coniosternum)) (Fig. 3). The species was described based on 1 male. Holotype: “Sogon-gomba – r. I-chyu, ver[khov’ya] Goluboi [reki], kon[ets] VII.[19]00 (Kozlov”)” [in Cyrillic, = Sogon-gomba, I-Chu River, upper reaches of Golubaya Rover, end of VII.[19]00 (Kozlov)], “Tibet, Kozl[ov] VII.[19]00,” “infumatum Beck[er],” “5,” “Holotypus ♂ Coniosternum infumatum Becker, 1907: 256 det. A.L. Ozerov, 2010,” designated by A.L. Ozerov (Ozerov and Krivosheina, 2011). The inventory number is INS_DIP_0001153. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; the left wing is strongly fractured at its base. The current name is Scathophaga infumata (Becker, 1907) (Gorodkov, 1986).

Fig. 3.

Coniosternum infumatum Becker, 1907, holotype. Scale bar 1 mm.

fulvisetis Becker, 1915 (Becker, 1915 : 64 (Pleurochaeta)) (Fig. 4). The species was described based on 1 male. Holotype: golden circle, “Polyarnyi Ural, s[ever] Tobol’s[koi] gub[ernii] 6.VII.[19]09 (F. Zaitsev)” [in Cyrillic, = Polar Urals, north of Tobolsk Governorate, 6.VII.[19]09 (F. Zaitsev)], “Pleurochaeta fulvisetis B. det. Becker,” “Holotypus ♂ Pleurochaeta fulvisetis Becker, 1915: 64 det. A.L. Ozerov, 2012,” “Pleurochaetella simplicipes (Becker) det. A.L. Ozerov, 2012.” The inventory number is INS_DIP_0001154. The holotype is missing the 2nd–5th tarsomeres in the right middle leg. The current name is Pleurochaetella simplicipes (Becker, 1900) (Gorodkov, 1986).

Fig. 4.

Pleurochaeta fulvisetis Becker, 1915, holotype. Scale bar 1 mm.

longinqua Becker, 1915 (Becker, 1915 : 66 (Scopeuma)) (Fig. 5). The species was described based on 4 specimens. Lectotype, male: “Karskaya tundra, s[ever] Tobol’s[koi] gub[ernii] 12.VII.[19]09 (F Zaitsev)” [in Cyrillic, = Kara Tundra, north of Tobolsk Governorate, 12.VII.[19]09 (F. Zaitsev)], “Lectotypus, ♂ Scopeuma longinqua Becker, 1915: 66 des. by A.L. Ozerov, 2012,” “Scathophaga cordylurina (Holmgren) det. A.L. Ozerov, 2012.” The inventory number is INS_DIP_0001155. The abdomen is cut off and placed together with the genitalia in a vial pinned under the specimen; the genitalia have lost color. Otherwise the lectotype is well preserved.

Fig. 5.

Scopeuma longinqua Becker, 1915, lectotype. Scale bar 1 mm.

Paralectotypes, 2 females, same locality. One more paralectotype, labeled similarly to the lectotype, is probably deposited at MNB in Berlin. The current name is Scathophaga cordylurina (Holmgren, 1883) (Gorodkov, 1986).

obscuripes Becker, 1915 (Becker, 1915 : 64 (Acerocnema)) (Fig. 6). The species was described based on 1 male. Holotype: golden circle, “Karskaya tundra, s[ever] Tobol’s[koi] gub[ernii] 12.VII.[19]09 (F Zaitsev)” [in Cyrillic, = Kara Tundra, north of Tobolsk Governorate, 12.VII.[19]09 (F. Zaitsev)], “Acerocnema Beck[er],” “obscuripes Beck[er],” “Acerocnema obscuripes B. det. Becker,” “Holotypus ♂ Acerocnema obscuripes Becker, 1915: 64 det. A.L. Ozerov, 2012,” “Nanna obscuripes (Becker) det. A.L. Ozerov, 2012.” The inventory number is INS_DIP_0001156. The left wing is partly damaged; otherwise the holotype is well preserved. The current name is Cleigastra obscuripes (Becker, 1915) (Ozerov and Krivosheina, 2023).

Fig. 6.

Acerocnema obscuripes Becker, 1915, holotype. Scale bar 1 mm.

orbitalis Becker, 1915 (Becker, 1915 : 65 (Scopeuma)) (Fig. 7). The species was described based on 1 female. Holotype: golden circle, “Pe-mal, s[ever] Tobol’s[koi] gub[ernii] 15.VIII.[19]09 (F. Zaitsev)” [in Cyrillic, = Pe-Mal, north of Tobolsk Governorate, 15.VIII.[19]09 (F. Zaitsev)], “orbitalis Beck[er],” “Scatophaga orbitalis B. det. Becker,” “Holotypus, Scopeuma orbitalis Becker, 1915: 65 det. A.L. Ozerov, 2012,” “Scathophaga nigripalpis (Becker [1907]) det. A.L. Ozerov, 2012.” The inventory number is INS_DIP_0001157. The holotype is well preserved. The current name is Scathophaga nigripalpis (Becker, 1907) (Gorodkov, 1986).

Fig. 7.

Scopeuma orbitalis Becker, 1915, holotype. Scale bar 1 mm.

perfecta Becker, 1907 (Becker, 1907 : 3 (Scatophaga)) (Fig. 8). The species was described based on 1 male. Holotype: golden circle, “na tsvetakh” [in Cyrillic, = on flowers], “Prijansky Tundra zw[ischen] Lena und Jana Brusnew 8, 9.VI.1902,” “Scatophaga perfecta Beck. det. Becker,” “Holotypus Scatophaga perfecta Becker, 1907: 3 det. A.L. Ozerov, 2012,” “Scatophaga apicalis Curtis ♂ Gorodkov det. [19]73.” The inventory number is INS_DIP_0001158. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen. The left fore leg is detached and glued onto a separate card; the right wing is partly damaged. The current name is Scathophaga apicalis (Curtis in Ross, 1835) (Gorodkov, 1986).

Fig. 8.

Scathophaga perfecta Becker, 1907, holotype. Scale bar 1 mm.

septentrionalis Becker, 1897 (Becker, 1897 : 397 (Scatophaga)) (Fig. 9). The species was described based on 5 specimens. Lectotype, male: “stoyanka u g[ory] Chernysheva. Nov[aya] Zem[lya] 5.VIII.[18]96 (Yakobson)” [in Cyrillic, = camp near Chernysheva Mt., Novaya Zemlya, 5.VIII.[18]96, Jacobson], “Scatophaga septentrionalis Becker,” “Lectotypus, ♂ Scatophaga septentrionalis Becker, 1897: 397 designation by A.L. Ozerov, 2011,” “Scatophaga varipes (Holmgren, [1883]) ♂ Gorodkov det. 1972.” The inventory number is INS_DIP_0001159. Both wings are partly damaged; otherwise the lectotype is well preserved.

Fig. 9.

Scathophaga septentrionalis Becker, 1897, lectotype. Scale bar 1 mm.

Paralectotypes, 2 males labeled similarly to the lectotype. Two paralectotypes identified by J.R. Vockeroth as Scathophaga varipes (Holmgren, 1883) are deposited at MNB (Ozerov and Krivosheina, 2013). The current name is Scathophaga varipes (Holmgren, 1883) (Gorodkov, 1986).

varitibia Becker, 1897 (Becker, 1897 : 400 (Microprosopa)) (Fig. 10). The species was described based on a single specimen, which Th. Becker considered a male. Later, A.L. Ozerov (Ozerov and Krivosheina, 2013) examined this specimen, found it to be a female, and synonymized the name with Allomyella frigida (Holmgren, 1883). Holotype, female: “Mal[ye] Karmakuly, Novaya Zemlya 16.VII.[18]96 (Yakobson)” [in Cyrillic, = Malye Karmakuly, Novaya Zemlya, 16.VII.[18]96, Jacobson], “Microprosopa varitibia B[ecker],” “Holotypus ♀ Microprosopa varitibia Becker, 1897: 400 det. A.L. Ozerov, 2011.” The inventory number is INS_DIP_0001160. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; the genitalia have lost color. The left fore leg and the right middle and hind legs are missing; both wings are partly damaged. The current name is Microprosopa frigida (Holmgren, 1883) (Ozerov and Krivosheina, 2023).

Fig. 10.

Microprosopa varitibia Becker, 1897, holotype. Scale bar 1 mm.

Species Described by I.A. Portschinsky

amplipennis Portschinsky, 1887 (Portschinsky, 1887 : 199 (Scatophaga)) (Fig. 11). The species was described based on males and females, but the exact number of specimens was not specified. Lectotype, male: “Scatophaga amplipennis” (handwritten by I.A. Portschinsky), “Lectotypus, ♂ Scatophaga amplipennis Portschinskiy, 1887: 199 ces. by Gorodkov, 1967: 446 cet. A.L. Ozerov, 2011,” designated by K.B. Gorodkov (1967 : 446). According to Portschinsky (1887 : 200), “the species was discovered during Mr. Przewalski’s expedition, high in the upper reaches of the Yellow River in Northeastern Tibet.” The upper reaches of the Yellow River [= Huang He] lie at altitudes greater than 3000 m above sea level. The date and collector were not specified. The inventory number is INS_DIP_0001161. The lectotype is well preserved.

Fig. 11.

Scathophaga amplipennis Portschinsky, 1887, lectotype. Scale bar 1 mm.

Paralectotype, male labeled similarly to the lectotype.

Note. The label of the lectotype made by A.L. Ozerov contains misprints cet. and ces., which should read det. and des., respectively.

magnipennis Portschinsky, 1887 (Portschinsky, 1887 : 198 (Scatophaga)) (Fig. 12). The species was described based on males and females, but the exact number of specimens was not specified. Lectotype, male: “San-Kul” [in Cyrillic], “Scatophaga magnipennis” (handwritten by I.A. Portschinsky), “Lectotypus, ♂ Scatophaga magnipennis Portschinskiy, 1887: 199 ces. by Gorodkov, 1967: 445 cet. A.L. Ozerov, 2011,” designated by K.B. Gorodkov (1967 : 445). According to Portschinsky (1887 : 200), “the species was discovered during Mr. Przewalski’s expedition, high in the upper reaches of the Yellow River in Northeastern Tibet” (see comments to the preceding species). The date and collector were not specified. The inventory number is INS_DIP_0001162. The lectotype is well preserved.

Fig. 12.

Scathophaga magnipennis Portschinsky, 1887, lectotype. Scale bar 1 mm.

Paralectotype, female labeled as the lectotype, but with the locality spelled as “Son-Kul” [in Cyrillic].

Note. The label of the lectotype made by A.L. Ozerov contains misprints cet. and ces., which should read det. and des., respectively.

Species Described by A.A. Stackelberg

paradoxopyga Stackelberg, 1952 (Stackelberg, 1952 : 405 (Acerocnema)) (Fig. 13). The species was described based on 3 males; the holotype was not explicitly designated by A.A. Stackelberg, but “Typ” is written on the identification label of one male. Holotype, male: golden circle, “Nov[aya] Zemlya, Matochkin Shar, okr. p. g. o. 11.VII.[1]925 (Vakulenko)” [in Cyrillic, = Novaya Zemlya, Matochkin Shar, env. of Polar Geophysical Observatory, 11.VII.[1]925, Vakulenko], “Acerocnema paradoxopyga Typ. sp. n. [19]41 Stackelberg det.,” “Holotypus Acerocnema paradoxopyga Stackelberg det. A.L. Ozerov, 2013.” The inventory number is INS_DIP_0001163. The holotype is glued onto a pinned card. The right middle and hind legs, the left hind tibia and tarsus are missing; the right wing is partly damaged.

Fig. 13.

Acerocnema paradoxopyga Stackelberg, 1952, holotype. Scale bar 1 mm.

Paratype, male: “o-v Vrangelya, Aternon–Rodzhers 27.VI.[1]939 (Portenko)” [in Cyrillic, = Wrangel Island, Aternon–Rogers, 27.VI.[1]939, Portenko]. The label refers to the area between the Rogers Bay and the Aternon Mountain, 10 km west of Ushakovskoe on the Rogers Bay coast. Another paratype, a male labeled similarly to the holotype, could not be located.

portenkoi Stackelberg, 1952 (Stackelberg, 1952 : 406 (Microprosopa)) (Fig. 14). The species was described based on 1 male. Holotype: golden circle, “o-v Vrangelya, Aternon–Rodzhers 27.VI.[1]939 (Portenko)” [in Cyrillic, = Wrangel Island, Aternon–Rogers, 27.VI.[1]939, Portenko] [area between Rogers Bay and Aternon Mountain, 10 km West of Ushakovskoe on Rogers Bay coast], “Microprosopa portenkoi sp. n. ♂ Typ. Stackelberg det.,” “Holotypus ♂ Microprosopa portenkoi Stackelberg, 1952 : 406 det. A.L. Ozerov, 2011.” The inventory number is INS_DIP_0001164. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; the genitalia have lost color. The left hind leg is missing; only the 1st tarsomere is preserved in the right hind tarsus.

Fig. 14.

Microprosopa portenkoi Stackelberg, 1952, holotype. Scale bar 1 mm.

Species Described by K.B. Gorodkov

sagittifera Gorodkov, 1974 (Gorodkov, 1974 : 388 (Cordilura)) (Fig. 15). The species was described based on 9 males and 8 females. Holotype, male: “Khargun, 6 km O Kyren, Sayany, osok[ovoe] boloto 11.VII.[1]965 (Gorodkov)” [in Cyrillic, = Buryatia, Khargun, 6 km E of Kyren, Sayan, sedge bog, 11.VII.[1]965, Gorodkov], “700 m,” “Holotypus ♂ Cordilura sagittifera Gorodkov, 1974: 388 cet. A.L. Ozerov, 2011.” The inventory number is INS_DIP_0001165. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; the right wing is partly damaged.

Fig. 15.

Cordilura sagittifera Gorodkov, 1974, holotype. Scale bar 1 mm.

Note. The label of the holotype made by A.L. Ozerov contains misprints cet. and ces., which should read det. and des., respectively.

Paratypes: 6 males and 8 females labeled similarly to the holotype; a vial with genitalia pinned under one specimen; 1 male: “Env. of Irkutsk, 4.VI.[1]911 (Malashedova)” [in Cyrillic], a vial with genitalia pinned under the specimen; 1 male: “Lake Tsagan, 100 verst W of Urga, Mongolia, 28–29.VI.[1]924, (P. Kozlov)” [in Cyrillic], a vial with genitalia pinned under the specimen.

sibirica Gorodkov, 1974 (Gorodkov, 1974 : 388 (Cordilura)) (Fig. 16). The species was described based on 1 male. Holotype: “[Buryatiya] Khargun, 6 km O Kyren, Sayany, osok[ovoe] boloto 11.VII.[1]965 (Gorodkov)” [in Cyrillic, = Buryatia, Khargun, 6 km E of Kyren, Sayan, sedge bog, 11.VII.[1]965, Gorodkov], “700 m,” “Holotypus ♂ Cordilura sibirica Gorodkov, 1974 : 388 cet. A.L. Ozerov, 2011.” The inventory number is INS_DIP_0001166. The tip of the abdomen is cut off and placed together with the genitalia in a vial pinned under the specimen.

Fig. 16.

Cordilura sibirica Gorodkov, 1974, holotype. Scale bar 1 mm.

Note. The label of the holotype made by A.L. Ozerov contains misprints cet. and ces., which should read det. and des., respectively.

stackelbergi Gorodkov, 1967 (Gorodkov, 1967 : 448 (Okeniella)) (Fig. 17). The species was described based on 16 males and 7 females. Holotype, male: “okr[estnosti] Mondy, Sayany, Tunkinskie gol’tsy 22.VII.[1]965 (Gorodkov)” [in Cyrillic, = env. of Mondy, Sayan Mts., Tunkinskie Goltsy Range, 22.VII.[1]965, Gorodkov], “r[eka] Khulugaima, 2500 m, golets, boloto” [in Cyrillic, = Khulugaima [Khulugaisha] River, 2500 m, bald mountain, bog], “Holotypus, ♂ Okeniella stackelbergi Gorodkov, 1967.” The inventory number is INS_DIP_0001167. The holotype is double mounted with a minuten pin and a card. The specimen is well preserved.

Fig. 17.

Okeniella stackelbergi Gorodkov, 1967, holotype. Scale bar 1 mm.

Paratypes: 2 males and 2 females labeled similarly to the holotype; 1 female, same locality but 27.VII.[1]965; 8 males and 7 females: “env. of Mondy, Sayan Mts., Tunkinskie Goltsy Range, 28.VII.[1]965 (Gorodkov)” [in Cyrillic], “upper reaches of Ikhe-Ukhgun River, bald mountain belt, bog, 2200 m” [in Cyrillic]. Gorodkov (1967) mentioned 7 females, but examination of the collection yielded 10 specimens. The remaining paratypes (5 males) could not be located. The current name is Pogonota stackelbergi (Gorodkov, 1967) (Ozerov and Krivosheina, 2023).

zaitzevi Gorodkov, 1974 (Gorodkov, 1974 : 386 (Cordilura)) (Fig. 18). The species was described based on 11 males and 6 females. Holotype, male: “Mongoliya, Tsentral’nyi aimak, Songino, poima i stepnye sklony 3–4.IX.[1]969 (Zaitsev)” [in Cyrillic, = Mongolia, Central Aimag, Songino, floodplain and steppe slopes, 3– 4.IX.[1]969 (Zaitsev)], “zaitzevi sp. n. Gorodkov det.,” “Holotypus ♂ Cordilura zaitzevi Gorodkov, 1974: 386 cet. A.L. Ozerov, 2011.” The inventory number is INS_DIP_0001168. The tip of the abdomen is cut off and placed together with the genitalia in a vial pinned under the specimen.

Fig. 18.

Cordilura zaitzevi Gorodkov, 1974, holotype. Scale bar 1 mm.

Note. The label of the holotype made by A.L. Ozerov contains misprints cet. and ces., which should read det. and des., respectively.

Paratypes: 3 females labeled similarly to the holotype; 3 males: “Mongoliya, Tsentr[al’nyi] Aimak, Songino 7.VII.[19]70 (Narchuk)” [in Cyrillic, = Mongolia, Central Aimag, Songino, 7.VII.[19]70, Nartshuk]; 2 males: “Mongoliya, Khent[eiskii] aimak, r[eka] Kerulen, 40 km ZYuZ Bayan-Obo 28.VII.[1]971 (Kerzhner)” [in Cyrillic, = Mongolia, Khentii Aimag, 40 km WSW of Bayan-Obo, 28.VII.[1]971, Kerzhner], with the genitalia on a card pinned under one specimen; 2 females: “[Buryatiya] Khargun, 6 km O Kyren, Sayany, osok[ovoe] boloto 11.VII.[1]965 (Gorodkov)” [in Cyrillic, = Buryatia, Khargun, 6 km E of Kyren, Sayan Mts., sedge bog, 11.VII.[1]965, Gorodkov], “700 m”; 1 male: “[Yakutiya] Nikol’skaya slobodka, Namsk[oi] ul[us] 26.VI.[19]01 (Gerts)” [in Cyrillic, = Yakutia, Nikolskaya Slobodka, Namskoi Ulus, 26.VI.[19]01, Herz]; 1 male: “Malokrasnoyarsk na Irtyshe, n[izhnee] Semir[ech’e] 27.VII.[1]926 (Vereshchagin)” [in Cyrillic, = Kazakhstan, East Kazakhstan Province, Malokrasnoyarsk on Irtysh River, Lower Semirechye, 27.VII.[1]926, Vereshchagin] (K.B. Gorodkov erroneously cited the date as 21.VII); 1 male: “Yakutsk 30.VI.[1]927 (Moskvin)” [in Cyrillic], with the genitalia on a card pinned under the specimen; 1 female: “Yakutsk 30.VII.[1]927 (Moskvin)” [in Cyrillic]; 1 male: “Yakutsk 8.VIII.[1]927 (Moskvin)” [in Cyrillic]; 1 male: “Novopokrovskoe, pr[avyi] b[ereg] Amgi, Yakut[iya] 22.VII.[1]928 (Pripuzov)” [in Cyrillic, = Novopokrovskoe, right bank of Amga, Yakutia, 22.VII.[1]928 (Pripuzov)].

Species Described by A.L. Ozerov

bicoloripes Ozerov, 1997 (Ozerov, 1997 : 1424 (Cordilura (Cordilurina)). The species was described based on 13 males and 14 females. Holotype and paratypes, 11 males and 13 females (ZMMSU). Two paratypes were gifted to the ZIN collection by A.L. Ozerov: 1 male: “Yuzh[noe] Primor’e, 32 km YuV Ussuriiska 30.VIII.1987 (A. Antropov)” [in Cyrillic, = South Primorye, 32 km SE of Ussuriisk, 30.VIII.1987, A. Antropov]; 1 female: “Yuzh[noe] Primor’e, Kamenushka 6.VIII.1987 (A. Shatalkin)” [in Cyrillic, = South Primorye, Kamenushka, 6.VIII.1987, A. Shatalkin].

caucasica Ozerov, 1993 (Ozerov, 1993 : 68 (Norellia (Norelliosoma)). The species was described based on 33 males and 13 females. Holotype and paratypes, 30 males and 11 females (ZMMSU). Four paratypes were gifted to the ZIN collection by A.L. Ozerov: 1 male: “Kavkaz, Sev[ernaya] Osetiya, khr[ebet] Bakhty, Lapparyrag, 10 km yu[go]-v[ostochnee] Alagira, 1600 m 29.VI.1990 (A.L. Ozerov)” [in Cyrillic, = Caucasus, North Ossetia, Bakhty Range, Lapparyrag, 10 km SE of Alagir, 1600 m, 29.VI.1990, A.L. Ozerov] and 1 female, same locality, 28.VI.1990; 1 male: “Sev[ernaya] Osetiya, Alagir 29.VI.1990 (A. Shatalkin)” [in Cyrillic, = North Ossetia, Alagir, 29.VI.1990, A. Shatalkin], “Lapparyrag 1700 m” [in Cyrillic] and 1 female, same locality, 28.VI.1990. The current name is Norellisoma caucasicum (Ozerov, 1993) (Ozerov and Krivosheina, 2023).

nigricorpus Ozerov, 2019 (Ozerov, 2019 : 206 (Gimnomera)) (Fig. 19). The species was described based on 4 males and 2 females. Holotype, male: “pos[elok] Agapa, [r.] Pyasina, Taimyr 14.VII.[1]967 (Gorodkov)” [in Cyrillic, = Agapa Village, Pyasina River, Taimyr, 14.VII.[1]967, Gorodkov], “lug v doline ruch’ya” [in Cyrillic, = meadow in spring valley], “Holotypus, ♂, Gimnomera nigricorpus sp. n. det. A.L. Ozerov, 2017.” The inventory number is INS_DIP_0001169. The holotype is double mounted with a minuten pin and a card. The abdomen is cut off and placed together with the genitalia in a vial pinned under the specimen.

Fig. 19.

Gimnomera nigricorpus Ozerov, 2019, holotype. Scale bar 1 mm.

Paratype, female: “Shmidt Chukotskii nats[ional’nyi] okr[ug] 11.VII.[1]971 (Gorodkov)” [in Cyrillic, = Cape Schmidt, Chukotka National Okrug, 11.VII.[1]971, Gorodkov], “pyatnistaya tundra” [in Cyrillic, = patchy tundra]. Other paratypes, 3 males and 1 female, are deposited in the ZMMSU collection.

nudipedis Ozerov, 2019 (Ozerov, 2019 : 207 (Gimnomera)) (Fig. 20). The species was described based on 3 males. Holotype: “pos[elok] Agapa, [r.] Pyasina, Taimyr 14.VII.[1]967 (Gorodkov)” [in Cyrillic, = Agapa Village, Pyasina River, Taimyr, 14.VII.[1]967, Gorodkov], “ivnyak, dolina ruch’ya” [in Cyrillic, = willow shrubs in spring valley], “Holotypus, ♂, Gimnomera nudipedis sp. n. det. A.L. Ozerov, 2018.” The inventory number is INS_DIP_0001170. The holotype is double mounted with a minuten pin and a card. The abdomen is cut off and placed together with the genitalia in a vial pinned under the specimen. All tarsomeres except the 1st are missing in the right hind leg.

Fig. 20.

Gimnomera nudipedis Ozerov, 2019, holotype. Scale bar 1 mm.

Paratypes: 1 male: “Tiksi, Yakutiya 10 km k yu[gu] 9.VII.[1]957 (Gorodkov)” [in Cyrillic, = Tiksi, Yakutia, 10 km to S, 9.VII.[1]957, Gorodkov], “dolina ruch’ya s ivoi” [in Cyrillic, = spring valley with willows], with genitalia in a vial pinned under the specimen; 1 male is deposited in the ZMMSU collection.

palliseta Ozerov, 2019 (Ozerov, 2019 : 208 (Gimnomera)). The species was described based on 6 males and 7 females. The holotype and paratypes (4 males and 7 females) are deposited in the ZMMSU collection.

Paratypes: 1 male: “[Krasnoyarsk Territory], Dudinka, Yenisei 3.VII.[1]967 (Gorodkov)” [in Cyrillic], “poima Eniseya, ol’shanik” [in Cyrillic, = Yenisei floodplain, alder forest]; 1 male: “Ust’-Kut na r[eke] Lene, Irkut[skaya] obl[ast’] 8.VI.[1]957 (Gorodkov)” [in Cyrillic, = Ust’-Kut on Lena River, Irkutsk Province, 8.VI.[1]957, Gorodkov], “syroi les” [in Cyrillic, = humid forest].

richterae Ozerov, 2015 (Ozerov, 2015 : 211 (Acerocnema)) (Fig. 21). The species was described based on 1 male. Holotype: “Khamnei, Buryatiya, poima r[eki] Shibertui 29.VI.1971 (V. Rikhter)” [in Cyrillic, = Khamnei, Buryatia, Shibertui River floodplain, 29.VI.1971, V. Richter], “Holotypus, ♂, Acerocnema richterae sp. n. det. A.L. Ozerov, 2015.” The inventory number is INS_DIP_0001171. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; otherwise the specimen is well preserved.

Fig. 21.

Acerocnema richterae Ozerov, 2015, holotype. Scale bar 1 mm.

taimyrica Ozerov, 2017 (Ozerov, 2017 : 93 (Microprosopa (Allomyella))) (Fig. 22). The species was described based on 1 male and 1 female. Holotype, male: “oz[ero] Engel’gardt, SV bereg, Taimyr 2.VIII.[1]967 (Gorodkov)” [in Cyrillic, = Krasnoyarsk Territory, Lake Engelgardt, NE shore, Taimyr, 2.VIII.[1]967, Gorodkov], “bereg ozera v tundre” [in Cyrillic, = lake shore in tundra], “Holotypus, ♂, Microprosopa taimyrica sp. n. det. A.L. Ozerov, 2016.” The inventory number is INS_DIP_0001172. The holotype is double mounted with a minuten pin and a card. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; both wings are partly damaged.

Paratype: female, labeled similarly to the holotype (ZMMSU).

Fig. 22.

Microprosopa (Allomyella) taimyrica Ozerov, 2017, holotype. Scale bar 1 mm.

Species Described by A.L. Ozerov and M.G. Krivosheina

alashanica Ozerov et Krivosheina, 2019 (Ozerov and Krivosheina, 2019 : 106 (Scathophaga)) (Fig. 23). The species was described based on 2 males and 2 females. Holotype, male: “Dyn-yuan’-in, s[evernaya] Alashan’ 30.V.[1]908 (Kozlov)” [in Cyrillic, = Dyn Yuan In, Northern Alashan, 30.V.[1]908, Kozlov], “Holotypus, ♂, Scathophaga alashanica sp. n. det. A.L. Ozerov, 2018.” The inventory number is INS_DIP_0001173. The first abdominal sternite in the holotype is partly damaged; the tip of the abdomen is cut off, but the corresponding genitalia preparation could not be located; otherwise the specimen is well preserved.

Fig. 23.

Scathophaga alashanica Ozerov et Krivosheina, 2019, holotype. Scale bar 1 mm.

Paratypes: 1 male and 2 females, labeled similarly to the holotype, but with dates 29, 30.V.[1]908 (Kozlov). The genitalia and remains of the abdomen are placed in a vial pinned under the male specimen.

buryatica Ozerov et Krivosheina, 2019 (Ozerov and Krivosheina, 2019 : 102 (Scathophaga)) (Fig. 24). The species was described based on 2 males. Holotype: “Samarta, V[ostochnye] Sayany, ok[olo] 40 km NO [pos.] Mondy 1.VIII.[1]965 (Gorodkov)” [in Cyrillic, = Samarta, East Sayan, ca 40 km NE of Mondy, 1.VIII.[1]965, Gorodkov], “v poselke” [in Cyrillic, = in settlement], “Holotypus, ♂, Scathophaga buryatica sp. n. det. A.L. Ozerov, 2018.” The inventory number is INS_DIP_0001174. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen.

Paratype: male, deposited in the ZMMSU collection.

Fig. 24.

Scathophaga buryatica Ozerov et Krivosheina, 2019, holotype. Scale bar 1 mm.

incompleta Ozerov et Krivosheina, 2020 (Ozerov and Krivosheina, 2020 : 228 (Scathophaga)). The species was described based on 21 males and 9 females. Holotype: male (ZMMSU).

Paratypes: 1 male: “Krym, Krymskii prirodnyi zapovednik 24.VII.1936 (V. Geptner)” [in Cyrillic, = Crimea, Crimean Nature Reserve, 24.VII.1936, V. Geptner]; 2 males: “Azerbaidzhan, vodokhranilishche Dzheiranbatan 17.X.1985 (Narchuk)” [in Cyrillic, = Azerbaijan, Ceyranbatan Reservoir, 17.X.1985, Nartshuk] (provisionally deposited in the ZMMSU collection).

gorodkovi Ozerov et Krivosheina, 2018 (Ozerov and Krivosheina, 2018 : 210 (Acerocnema)) (Fig. 25). The species was described based on 1 male. Holotype: “okr[estnosti] Khabarovska, Malyshevo, ok[olo] Petropavlovki 13.V.[1]973 (Gorodkov)” [in Cyrillic, = environs of Khabarovsk, Malyshevo, near Petropavlovka, 13.V.[1]973, Gorodkov], “shirokolistvennyi les” [in Cyrillic, = broad-leaved forest], “Holotypus, ♂, Acerocnema gorodkovi sp. n. det. A.L. Ozerov, 2018.” The inventory number is INS_DIP_0001175. The holotype is double mounted with a minuten pin and a card. The abdomen is cut off and placed together with the genitalia in a vial pinned together with the specimen.

Fig. 25.

Acerocnema gorodkovi Ozerov et Krivosheina, 2018, holotype. Scale bar 1 mm.

grunini Ozerov et Krivosheina, 2017 (Ozerov and Krivosheina, 2017 : 2 (Cordilura)) (Fig. 26). The species was described based on 1 male. Holotype: “Dol[ina] r[eki] Sankhobe, Primorsk[aya] obl[ast’] 17.V.[1]937 (Grunin)” [in Cyrillic, = Sankhobe River valley, Primorskaya Province, 17.V.[1]937, Grunin], “Holotypus, ♂, Cordilura grunini sp. n. det. A.L. Ozerov, 2017.” The inventory number is INS_DIP_0001176. The holotype is double mounted with a minuten pin and a card. The abdomen is cut off and placed together with the genitalia in a vial pinned together with the specimen. Both wings and the right middle leg are missing; the tarsus and the distal part of the tibia are missing in the left hind leg.

Fig. 26.

Cordilura grunini Ozerov et Krivosheina, 2017, holotype. Scale bar 1 mm.

maritima Ozerov et Krivosheina, 2016 (Ozerov and Krivosheina, 2016 : 98 (Cleigastra (Orthacheta))) (Fig. 27). The species was described based on 1 male and 1 female. Holotype, male: “Gribnoe, 10 km YuV Chernigovki, yu[zhnoe] Primor’e 19.V.1979 (A. Zinov’ev)” [in Cyrillic, = Gribnoe, 10 km SE of Chernigovka, South Primorye, 19.V.1979, A. Zinoviev], “Holotypus, ♂, Cleigastra (Orthacheta) maritima sp. n. det. A.L. Ozerov, 2015.” The inventory number is INS_DIP_0001177. The holotype is double mounted with a minuten pin and a card. The abdomen is cut off and placed together with the genitalia in a vial pinned to the specimen; otherwise the holotype is well preserved.

Paratype: male, deposited in the ZMMSU collection.

Fig. 27.

Cleigastra (Orthacheta) maritima Ozerov et Krivosheina, 2016, holotype. Scale bar 1 mm.

nartshukae Ozerov et Krivosheina, 2015 (Ozerov and Krivosheina, 2015 : 489 (Cordilura)) (Fig. 28). The species was described based on 1 female. Holotype: “verkh[ov’e] Chapigou, prit[ok] Shufana, yu[zhnoe] Primor’e 5.VII.[1]962 (Narchuk)” [in Cyrillic, = Upper reaches of Chapigou, tributary of Shufan River, South Primorye, 5.VII.[1]962, Nartshuk], “Holotypus, ♀ Cordilura nartshukae sp. n. det. A.L. Ozerov, 2015.” The inventory number is INS_DIP_0001178. The holotype is well preserved.

Fig. 28.

Cordilura nartshukae Ozerov et Krivosheina, 2015, holotype. Scale bar 1 mm.

negrobovi Ozerov et Krivosheina, 2017 (Ozerov and Krivosheina, 2017 : 4 (Cordilura)) (Fig. 29). The species was described based on 1 male. Holotype: “V[ostochnye] Sayany, Arshan, Tagyrkha 3.VII.[1]965 (Negrobov)” [in Cyrillic, = East Sayan, Arshan, Tagyrkha, 3.VII.[1]965, Negrobov], “boloto Negrobov 96” [in Cyrillic, = bog, Negrobov, 96], “Holotypus, ♂, Cordilura negrobovi sp. n. det. A.L. Ozerov, 2017.” The inventory number is INS_DIP_0001179. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned together with the specimen. The right middle leg is missing.

Fig. 29.

Cordilura negrobovi Ozerov et Krivosheina, 2017, holotype. Scale bar 1 mm.

nubilosa Ozerov et Krivosheina, 2020 (Ozerov and Krivosheina, 2020 : 461 (Cordilura)) (Fig. 30). The species was described based on 1 male. Holotype: “versh[ina] g[ory] Oblachnoi, yuzh[nyi] Sikhote-Alin’, Prim[orskii] kr[ai] 3.VII.[1]962 ([O.V.] Kovalev) v[ysota] 1856 [m]” [in Cyrillic, = Top of Oblachnaya Mt., South Sikhote-Alin, Primorskii Territory, 3.VII.1962, O.V. Kovalev, alt. 1856 m], “Holotypus, ♂ Cordilura nubilosa Ozerov & Krivosheina sp. n. det. A.L. Ozerov, 2020.” The inventory number is INS_DIP_0001180. The abdomen of the holotype is cut off and placed together with the genitalia in a vial pinned under the specimen; the left wing is torn.

Fig. 30.

Cordilura nubilosa Ozerov et Krivosheina, 2020, holotype. Scale bar 1 mm.

richterae Ozerov et Krivosheina, 2015 (Ozerov and Krivosheina, 2015 : 491 (Cordilura)) (Fig. 31). The species was described based on 2 males. Holotype: “Yu[zhnye] Kurily, Iturup, Rybakhi 5 km SW Kuril’ska 23.VI.1968 (V. Rikhter)” [in Cyrillic, = South Kuril Islands, Iturup, Rybakhi, 5 km SW of Kurilsk, 23.VI.1968, V. Richter], “Holotypus, ♂ Cordilura richterae sp. n. det. A.L. Ozerov, 2015.” The inventory number is INS_DIP_0001181. The right hind leg is missing; only the 1st tarsomere is preserved in the left fore tarsus.

Paratype: male, deposited in the ZMMSU collection.

Fig. 31.

Cordilura richterae Ozerov et Krivosheina, 2015, holotype. Scale bar 1 mm.