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Varanus rudicollis (GRAY, 1845)

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Higher TaxaVaranidae, Platynota, Varanoidea, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Roughneck Monitor
G: Rauhnackenwaran
Italian: Varano dal collo ruvido 
SynonymUaranus [sic] rudicollis GRAY 1845: 10
Varanus rudicollis — BOULENGER 1885: 313
Varanus rudicollis — DE ROOIJ 1915: 144
Varanus scutigerulus BARBOUR 1932 (fide TAYLOR 1963)
Varanus (Varanus) salvator scutigerulus — MERTENS 1942: 259
Varanus (Dendrovaranus) rudicollis — MERTENS 1942: 360
Varanus rudicollis — MERTENS 1950: 6
Varanus swarti MANGILI 1962: 30 (fide MERTENS 1964)
Varanus rudicollis — TAYLOR 1963: 915
Varanus rudicollis — DE LISLE 1996: 133
Varanus rudicollis — MANTHEY & GROSSMANN 1997: 291
Varanus (Empagusia) rudicollis — ZIEGLER & BÖHME 1997
Varanus rudicollis — GRISMER 2011
Varanus (Empagusia) rudicollis — KOCH et al. 2013
Varanus (Empagusia) rudicollis — BUCKLITSCH et al. 2016: 50 
DistributionS Burma (= Myanmar), S Thailand, W Malaysia (incl. Penang), Rhio Archipelago, Indonesia (Sumatra, Bangka, Borneo), Philippines ?

Type locality: Philippines (in error)  
Reproductionoviparous 
TypesHolotype: BMNH 1842.2.15.239
Holotype: MCZ 32264 [scutigerulus] 
Diagnosis 
CommentSynonymy: Mertens 1950 synonymized Varanus scutigerulus with V. rudicollis. 
References
  • Amer, Sayed A. M. and Yoshinori Kumazawa 2008. Timing of a mtDNA gene rearrangement and intercontinental dispersal of varanid lizards. Genes Genet. Syst. 83: 275–280 - get paper here
  • Auliya, M. 2006. Taxonomy, Life History, and conservation of giant reptiles in west Kalimantan. Natur und Tier Verlag, Münster, 432 pp. - get paper here
  • Ávalos, J. de & Martínez Carrión, P. 1997. Warane. Reptilia (Münster) 2 (5): 16-21 - get paper here
  • Barbour, Thomas 1932. A new Bornean monitor. Proc. New England zool. Club 13: 1-2
  • Bayless, Mark K. 1997. The rough-neck Monitor lizard (Varanus rudicollis). Bull. Chicago Herp. Soc. 32 (12): 250-252 - get paper here
  • Bennett, Daniel 1993. A review of some literature concerning the rough-necked monitor lizard Varanus rudicollis. Reptilian 1 (9): 7-10
  • Bennett, Daniel; Liat, Lim Boo 1995. A note on the distribution of Varanus dumerilii and V. rudicollis in Peninsular Malaysia. Malayan Nature Journal 49 (2):113-116.
  • Bochaton, Corentin; Martin Ivanov and Julien Claude 2019. Osteological criteria for the specific identification of Monitor lizards (Varanus Merrem, 1820) remains in subfossil deposits of Sundaland and continental Southeast Asia. Amphibia-Reptilia 40 (2): 219–232 - get paper here
  • Böhme,W. 2003. Checklist of the living monitor lizards of the world (family Varanidae). Zoologische Verhandelingen 341: 6-43 - get paper here
  • Bong Heang, Kiew 1987. An annotated checklist of the herpetofauna of Ulu Endau, Johore, Malaysia. Malayan Nature Journal 41 (2-3): 413-423.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • BUCKLITSCH, YANNICK; WOLFGANG BÖHME, ANDRÉ KOCH 2016. Scale Morphology and Micro-Structure of Monitor Lizards (Squamata: Varanidae: Varanus spp.) and their Allies: Implications for Systematics, Ecology, and Conservation. . Zootaxa 4153 (1): 001–192 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Cooper, Taylor; Amanda Liew, Gabriel Andrle, Elisabeth Cafritz, Hannah Dallas, Trent Niesen, Emily Slater, Joseph Stockert, Taylor Vold, Michael Young & Joseph Mendelson 2019. Latency in Problem Solving as Evidence for Learning in Varanid and Helodermatid Lizards, with Comments on Foraging Techniques. Copeia 107 (1): 78-84 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Gaulke, Maren 1991. Systematic relationships of the Philippine water monitors as compared with Varanus s. salvator, with a discussion of dispersal routes. Mertensiella 2: 154-167
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Chan K. Onn, Jesse L. Grismer, Perry L. Wood, Jr., and A. Norhayati 2010. A CHECKLIST OF THE HERPETOFAUNA OF THE BANJARAN BINTANG, PENINSULAR MALAYSIA. Russ. J. Herpetol. 17 (2): 147-160 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Hamilton, R. 2012. First Record of the Rough-necked Monitor Varanus rudicollis from Penang, Malaysia. Biawak 6 (2): 78 - get paper here
  • Horn,H.G. & Peters, G. 1982. Beiträge zur Biologie des Rauhnackenwarans, Varanus (Dendrovaranus) rudicollis Gray. Salamandra 18 (1-2): 29-40 - get paper here
  • Iannucci, Alessio; Marie Altmanová, Claudio Ciofi, Malcolm Ferguson-Smith, Massimo Milan, Jorge Claudio Pereira, James Pether, Ivan Rehák, Michail Rovatsos, Roscoe Stanyon, Petr Velenský, Petr Ráb, Lukáš Kratochvíl & Martina Johnson Pokorná 2019. Conserved sex chromosomes and karyotype evolution in monitor lizards (Varanidae). Heredity 123, 215–227 - get paper here
  • Koch A, Ziegler T, Böhme W, Arida E, Auliya M. 2013. Pressing Problems: Distribution, Threats, and Conservation Status of the Monitor Lizards (Varanidae: Varanus spp.) of Southeast Asia and the Indo-Australian Archipelago. Herp. Cons. Biol. 8 (Monograph 3): 1 − 62. - get paper here
  • Lisle, H.F. de 1996. Natural History of Monitor Lizards. Krieger, Malabar (Florida)
  • MANGILI 1962. . Giard. zool. Roma, 4: 30
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Mertens, R. 1950. Notes on some Indo-Australian monitors (Sauria, Varanidae). American Museum Novitates 1456: 1-7 - get paper here
  • Pianka, E. R. and D. R. King, eds. 2004. Varanoid Lizards of the World. Indiana University Press, 599 pp - get paper here
  • Pianka, Eric R. 2012. Can humans share spaceship earth? Amphibian & Reptile Conservation 6 (1): 1-24 - get paper here
  • Rese, R. 1988. Beitrag zur Lebenserwartung von Grosswaranen in Gefangenschaft. Sauria 10 (1): 25-26 - get paper here
  • Taylor, E.H. 1922. The lizards of the Philippine Islands. Department of Agriculture and Natural Resources, Bureau of Science, Government of the Philippine Islands, Manila, Publication no. 17: 269 pp. - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Werner, F. 1900. Reptilien und Batrachier aus Sumatra, gesammelt Herrn. Gustav Schneider jr., im Jahre 1897-1898. Zool. Jahrb. (syst) 13: 479-508 - get paper here
  • Ziegler, Thomas; Böhme, Wolfgang 1997. Genitalstrukturen und Paarungsbiologie bei squamaten Reptilien, speziell den Platynota, mit Bemerkungen zur Systematik. Mertensiella 8: 1-207
  • Zug, G.R. & Mulcahy, D.G. 2019. Identification guide Amphibians and reptiles of South Tanintharyi. Fauna & Flora International, 101 pp.
 
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